What was the attitude of leaders of the Ionian Greeks towards Persia?

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This probably could occupy a whole essay; pretty sure the only primary reference we have for the opinion of Ionian Greeks (i.e., Greeks literally from Ionia) is Herodotus. If you mean more broadly to include Athenians (who for many purposes are also considered "Ionian"), Athenian mercenaries (such as Xenophon) in the period after the Peloponnesian War had no problem with serving under a Persian.

Obviously, most Ionian leadership prior to the Persian War was anti-Persia (see eg, the Ionian Revolt: "The Persians found the Ionians difficult to rule."), which was a contributing factor to revolting against Persian rule. But, "medizing" was simultaneously something that most Greeks looked down upon but also seemed to have happened frequently enough to have its own term, so under the right circumstances different Ionians aligned themselves with Persia (it's been long enough since I read Herodotus, I can't remember if there are specific examples).

Furthermore, the attitude of all Greeks towards Persia seems to be opportunistic and changed over time. As an example, Sparta was vigorously anti-Persia during the Persian Wars, but 75 years later had no qualms about siding with Persia in the Peloponnesian Wars. As another example, Athenians saw no problems with serving in mercenaries for Persia after their ancestors had fought Persia a century prior.

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